Ai! Lá polin saca i quettar!
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A noun in The Etymologies of the 1930s glossed “rat” derived from primitive ᴹ✶nyadrō under the root ᴹ√NYAD “gnaw” (Ety/NYAD). Tolkien gave the intermediate form naðr, but it is not clear why the ð vanished; compare N. nadhor “pasture” < ᴹ√NAD (< ✱nadrō?), and indeed Tolkien had a variant archaic form naðor “rat” which shows the normal phonetic developments (EtyAC/NYAD).